Sambucus, breathing through the cork layer

Cork is formed by a cambium layer that itself almost always arises as secondary meristem from the collenchyma or parenchyma layer directly under the epidermis. Unlike sclerenchymal cells, collenchymal cells are still alive and have retained the ability to differentiate. Cork cambium cells only divide periclinally, creating the typical rows of daughter cells. In this way mainly cork cells are produced to the outside and to a much lesser extent (sometimes not at all) cork parenchyma is produced to the inside. Cork cambium, cork cells and cork parenchyma together are also called periderm. Developed cork cells are dead and their cell walls contain suberine, a greasy substance that repels water.